By default, all parts of your model tick at the same rate, as specified by the Update every... value in the run control dialogue window. However, you will sometimes want to get parts of the model updated less (or more) frequently than others. For example, you may have a model containing both trees and a crop. The crop you want to grow on a weekly basis, so you can capture its response to rainfall patterns, pest outbreaks etc. The trees grow slowly, and there is no point at all in calculating tiny increments on a week-by-week basis.